1The Faculty of Physical Culture, Sports and Tourism, Mari State University, Lenin Square 1, Yoshkar-Ola 424000, Russian Federation
2The Institute of Medicine and Natural Sciences, Mari State University, Lenin Square 1, Yoshkar-Ola 424000, Russian Federation
3Research Department, Federal State Organization, National Park Mari Chodra, Central Street 73, Krasnogorsk 425090, Mari El Republic, Russian Federation
4The Faculty of Biology, Tver State University, Zhelyabova 33, Tver 170100, Russian Federation