1Key Laboratory of Protection and Development Utilization of Tropical Crop Germplasm Resources, Ministry of Education, College of Environment and Plant Protection, Hainan University, Haikou, Hainan, 570228, P.R. China
2Department of Plant Biology and Pathology, School of Environmental and Biological Sciences, Rutgers, 59 Dudley Road, New Brunswick, New Jersey, 08901, USA