1Department of Fisheries and Aquaculture, Faculty of Agrotechnology and Food Science, Universiti Malaysia Terengganu, 21030, Kuala Terengganu, Malaysia
2Institute of Marine Biotechnology, Universiti Malaysia Terengganu, 21030, Kuala Terengganu, Malaysia
3Department of Biology, Dalhousie University, Halifax, NS, B3H 4R2, Canada