1Faculty of Agriculture and Life Science, Hirosaki University, 3 Bunkyo-cho, Hirosaki, Aomori 036-8561, Japan
2Graduate School of Bio-Applications and Systems Engineering, Tokyo university of agriculture and technology, Koganei, Tokyo 184-8588, Japan
3Institute for Environmental Management Technology, National Institute of Advanced Industrial Science and Technology, 16-1 Onogawa, Tsukuba, Ibaraki 305-0053, Japan