1Higher Agronomic Institute of Chott-Mariem, Sousse University, 4042-Chott Mariem, Tunisia
2UR13AGR09- Integrated Horticultural Production in the Tunisian Centre East, Regional Centre of Research on Horticulture and Organic Agriculture, University of Sousse, 4042, Chott-Mariem, Tunisia
3INRA, UMR1065 Santé et Agroécologie du Vignoble (SAVE), ISVV, F-33140 Villenave d’Ornon, France
4University of Bordeaux, Bordeaux Sciences Agro, ISVV, UMR1065 SAVE, F-33140 Villenave d’Ornon, France